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world of amoeboid organisms

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Class Tubulinea Smirnov et al., 2005 (=Lobosea Cavalier-Smith, 2004)

Diagnosis: Organisms producing lobose pseudopodia (lobopodia). The entire cell or individual pseudopodia (in polypodial cells) are tubular, cylindrical or sub-cylindrical, rounded in cross-section. If cells are flattened or branched they are capable of altering the locomotive form from a flattened, expanded one to monopodial or polypodial, with sub-cylindrical pseudopodia. Monoaxial flow of the cytoplasm in every pseudopodium or in the entire cell. No convincing evidence of ciliate stages. Two groups are testate, and two sorocarpic taxa are known. No sporocarpy has been reported.

  1. Order Elardia Kang et al. 2017
    Cells naked or covered with a hard test; tubular or produce tubular pseudopodia; if flattened or branched, capable of altering the locomotive form to monopodial or polypodial, with tubular pseudopodia. The least inclusive clade containing Amoeba proteus, Arcella intermedia and Rhizamoeba saxonica.

    1. Arcellinida Saville Kent, 1880
      Testate, inside an organic or mineral extracellular test of either self-secreted elements (calcareous, siliceous, or chitinoid) or recycled mineral particles bound together, with a single main opening.

    2. Euamoebida (Lepşi, 1960)
      Naked amoebae with subcylindrical pseudopodia in locomotion (or the entire cell is monopodial and subcylindrical); without alteration of the locomotive form to a flattened expanded and branched one; without adhesive uroid; glycocalyx amorphous, filamentous or consisting of prismatic, cup-shaped structures.

      1. Amoebina Cavalier-Smith, 2016
      2. Nolandina Cavalier-Smith, 2016

    3. Leptomyxida (Pussard and Pons, 1976) Page, 1987
      Naked amoebae with locomotive form altering from a flattened expanded or reticulate one to a subcylindrical monopodial one when in rapid movement or under specific conditions; adhesive uroidal structures always present.
      Flabellula, Gephyramoeba*, Leptomyxa, Rhizamoeba.

  2. Order Echinamoebida Cavalier-Smith, 2004
    Cells tubular, vermiform or flattened, with or without spine-like subpseudopodia; capable of adopting subcylindrical monopodial form under certain conditions.
    Echinamoeba, Micriamoeba, Vermamoeba.

  3. Order Corycida Kang et al. 2017
    Cells covered with flexible, leather-like coating forming one or several openings used to protrude pseudopodia or are enclosed in hard test made of spicules with multiple apertures.
    Amphizonella, Diplochlamys, Trichosphaerium.

  4. Order Trichosida Möbius, 1889
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